European equities traded in the US as American depositary receipts were higher late Tuesday morning, rising 0.59% to 1,393.03 on the S&P Europe Select ADR Index.

From continental Europe, the gainers were led by Spanish lender Banco Santander (SAN) and biotech company Evaxion Biotech (EVAX), which rose 3.8% and 2.5% respectively. They were followed by software firm SAP (SAP) and Spanish financial services company Banco Bilbao Vizcaya Argentaria (BBVA), which were up 2% and 1.9% respectively.

The decliners from continental Europe were led by semiconductor company Sequans Communications (SQNS) and accommodations booking company trivago (TRVG), which dropped 9.6% and 8.5% respectively. They were followed by biopharmaceutical companies Cellectis (CLLS) and argenx (ARGX), which were down 2.5% and 2% respectively.

From the UK and Ireland, the gainers were led by oil and gas company Shell (SHEL) and British bank Barclays (BCS), which increased 1.7% and 1.5% respectively. They were followed by medical device maker Smith & Nephew (SNN) and British financial services company HSBC (HSBC), which were up 1.4% and 1.1% respectively.

The decliners from the UK and Ireland were led by biopharmaceutical company Biodexa Pharmaceuticals (BDRX) and pharmaceutical company Silence Therapeutics (SLN), which fell 16% and 11% respectively. They were followed by biopharmaceutical companies Akari Therapeutics (AKTX) and Bicycle Therapeutics (BCYC), which lost 7.4% and 3.5% respectively.